Werner Conradie Publications

Peer Reviewed Publications

2025

Conradie W, Keates C, Greenbaum E, Lobón-Rovira J, Tolley KA, Benito M, Vaz Pinto P, van Breda RV, Verburgt L. 2025.  Systematics of African rough-scaled lizards, with description of two new species from eastern Angola (Squamata: Lacertidae: Ichnotropis Peters, 1854). Vertebrate Zoology 75: 627–672. https://doi.org/10.3897/vz.75.e167366

Tolley KA, Barends JM, Greenbaum E, Conradie W.  2025. A comprehensive phylogeny of night adders (Causus), African Journal of Herpetology. 74: 320–336. https://doi.org/10.1080/21564574.2025.2555826 

Barends JM, Stanton-Jones WK, Conradie W, Tolley KA. 2025. The herpetofauna of Meerkat National Park, Northern Cape province, South Africa. Koedoe 67: a1865. https://doi.org/10.4102/koedoe.v67i1.1865

Lobón-Rovira J, Vaz Pinto P. Cael G, Conradie W. 2025. Is morphological-based taxonomy still effective in modern times? A response to Parrinha et al. (2025). 61(2): 320--328. https://www.salamandra-journal.com/index.php/contents/2025-vol-61/2191-lob%C3%B3n-rovira,-j-,-p-vaz-pinto,-g-cael-w-conradie/file 

Lobón-Rovira J, Heinicke MP, Bauer AM,  Conradie W, Vaz Pinto P. 2025., Three New Endemic Species of Namib Day Geckos (Gekkonidae: Rhoptropus) From the Namibe Province, Angola. Ecology and Evolution, 15 (7): e71609. https://doi.org/10.1002/ece3.71609 

Benito M, Conradie W, Vaz Pinto P, Lobón-Rovira J (2025) A needle in a haystack: Rediscovery and revised description of Ichnotropis microlepidota Marx, 1956, from the central highlands of Angola. Zoosystematics and Evolution 101(2): 887-906. https://doi.org/10.3897/zse.101.136290

Lawson LP, Bittencourt-Silva GB, Conradie W, Portik DM, Loader SP (2025) A phylogenomic perspective reveals mitochondrial-nuclear discordance and previously undescribed species nested within a widespread East African Reed frog species (Hyperolius substriatus Ahl, 1931). PLoS ONE 20(4): e0318951. https://doi.org/10.1371/journal.pone.0318951 

Conradie W, Hundermark C, Kemp L, Keates C. 2025. New Pygmy Gecko (Goggia: Gekkonidae) from the arid Northern Cape Province of South Africa. Zootaxa, 5618 (4): 552–570. https://doi.org/10.11646/zootaxa.5618.4.5 

Lobón-Rovira J, Baptista NL, Clark T, Verburgt L, Jongsma GFM, Conradie W, Vermissimo L, Vaz Pinto, P. 2025. Filling the gaps: herpetological checklist of Mayombe National Park and Cabinda Province (Angola) shed light on one of the most unexplored corners of tropical Central Africa. African Journal of Herpetology, 1–59. https://doi.org/10.1080/21564574.2024.2421007

Cowan OS, Conradie W, Keates C, Telford NS, Petford MA, Egan V, Kolley KA. 2025. Lost lizards: The importance of targeted surveys in filling knowledge gaps for reptile conservation in South Africa. Journal for Nature Conservation, 84(126790). https://doi.org/10.1016/j.jnc.2024.126790. 

2024

Lobón-Rovira J, Marugán-Lobón J, Nebreda SM, Buckley D, Stanley EL, Glaw F, Conradie W,  Bauer AM 2024.  Adaptive or non-adaptive? Cranial evolution in a radiation of miniaturized day geckos. BMC Ecology and Evolution, 24(150). https://doi.org/10.1186/s12862-024-02344-w 

Kameni N MM, Tapondjou N WP, Conradie W. 2024. Phylogenetic placement of the Mount Bamboutos endemic skink, Trachylepis mekuana (Chirio and Ineich, 2001). Acta Herpetologica, 11(2):167–171. https://doi.org/10.36253/a_h-16188 

Keates C, Conradie W, S. Edwards S. 2024. Phylogeographic structuring in a widespread southern African grass snake (Psammophiinae: Psammophylax rhombeatus), African Journal of Herpetology, 73(2): 210–235. DOI: 10.1080/21564574.2024.2375496  

Angus O, Ping T, Telfrod N, Conradie W. 2024. A new species of Dainty frog (Anura: Pyxicephalidae:  Cacosternum) and the first endemic anuran to the Cederberg region of South Africa. Zootaxa, 5512 (1): 001–021. https://doi.org/10.11646/zootaxa.5512.1.1 

Keates C, Wasserman RJ., Conradie W, Dondofema F, Munyai L, Riddell E, Dalu F. 2024. Frogs of the Makuleke Contractual Park, northern Kruger National Park, Koedoe, 66(1): a1785. https://doi.org/10.4102/koedoe.v66i1.1785

Bayliss J, Bittencourt-Silva GB, Branch WR, Bruessow C, Collins S, Congdon TCE, Conradie W, Curran M, Daniels SR, Darbyshire I, Farooq H, Fishpool L, Grantham G, Magombo Z, Matimele H, Monadjem A, Monteiro J, Osborne J, Saunders J, Smith P, Spottiswoode CN, Taylor PJ, Timberlake J, Tolley KA, Érica Tovela E, Platts P. 2024. Recent biological surveys A biogeographical appraisal of the threatened South East Africa Montane Archipelago ecoregion. Scientific Reports, 14: 5971. https://doi.org/10.1038/s41598-024-54671-z 

Chaney T, Pauwels OSG, Nagy ZT, Gvoždík V, Kusamba C, Badjedjea G, Masudi FM, Akuboy JB, Ernst R, Trape J-F, Chirio L, Conradie W, Keates C, Wallach V, Zassi-Boulou A-G, Vaughan ER, Greenbaum E. 2024 Phylogenetics and Integrative Taxonomy of African Water Snakes (Squamata: Colubridae: Grayia). Herpetological Monographs, 38(1): 1–52. https://doi.org/10.1655/HERPMONOGRAPHS-D-23-00002.1  

2023

Conradie W, Lobón-Rovira J, Becker FS, Schmitz A, Vaz Pinto P. 2023. Flat gecko (Afroedura) diversity, endemism and speciation in the highlands and escarpments of Angola and Namibia. Namibian Journal of Environment. 8: 277–281

Becker FS, Baptista NL, Vaz Pinto P, Ernst R, Conradie W. 2023. The amphibians of the highlands and escarpments of Angola and Namibia. Namibian Journal of Environment. 8: 245–257

Lobón-Rovira J, Bauer AM, Vaz Pinto P, Trape J-F, Conradie W, Kusamba C, Júlio T, Cael G, Stanley EL, Hughes DF, Behangana M, Masudi FM, Pauwels OSG, Greenbaum E. 2023. Integrative revision of the Lygodactylus gutturalis (Bocage, 1873) complex unveils extensive cryptic diversity and traces its evolutionary history, Zoological Journal of the Linnean Society, 201(2): 447–492, https://doi.org/10.1093/zoolinnean/zlad123

Conradie W, Keates C, Verburgt L, Baptista NL, Harvey J. 2023. Contributions to the herpetofauna of the Angolan Okavango-Cuando- Zambezi River drainages. Part 3: Amphibians. Amphibian & Reptile Conservation 17(1 & 2) [General Section]: 19–56 (e325). http://amphibian-reptile-conservation.org/pdfs/Volume/Vol_17_nos_1-2/ARC_17_1-2_%5BGeneral_Section%5D_19-56_e325.pdf 

Rödel MO, Becker FS, Buiswalelo B, Conradie W, Channing A. 2023. Re-evaluation of the status of Bufo vertebralis grindleyi and Bufo jordani (Anura: Bufonidae). Salamandra 59(2): 143–157. https://www.salamandra-journal.com/index.php/home/contents/2023-vol-59/2117-roedel-m-o-f-s-becker-b-buiswalelo-w-conradie-a-channing

Reeves B & Conradie W. 2023 Breeding phenology and behaviour of the Endangered Hewitt’s ghost frog Heleophryne hewitti in the Baviaanskloof World Heritage Site, Eastern Cape Province, South Africa, Bioacoustics, https://doi.org/10.1080/09524622.2023.2193815 

Zhao Z, Conradie W, Pietersen DW, Jordaan A, Nicolau G, Edwards S, Riekert S, Heideman N. 2023. Diversification of the African legless skinks in the subfamily Acontinae (Family Scincidae). Molecular Phylogenetics and Evolution. 182:107747 https://doi.org/10.1016/j.ympev.2023.107747 

Premachandra T, Cauret CMS, Conradie W, Measey J, Evans BJ. 2023. Population genomics and subgenome evolution of the allotetraploid frog Xenopus laevis in southern Africa, G3 Genes|Genomes|Genetics, Volume 13, Issue 2, February 2023, jkac325, https://doi.org/10.1093/g3journal/jkac325

2022

Conradie W, Keates C, Verburgt L, Baptista NL, Harvey J, Júlio T, Neef G. 2022. Contributions to the herpetofauna of the Angolan Okavango- Cuando-Zambezi River drainages. Part 2: Lizards (Sauria), chelonians, and crocodiles. Amphibian & Reptile Conservation 16(2) [General Section]:181–214 (e322). http://amphibian-reptile-conservation.org/pdfs/Volume/Vol_16_no_2/ARC_16_2_[General_Section]_181-214_e322.pdf 

Conradie W, Schmitz A, Lobón-Rovira J, Becker FS, Vaz Pinto P, Hauptfleisch ML. 2022. Rock island melody remastered: two new species in the Afroedura bogerti Loveridge, 1944 group from Angola and Namibia. Zoosystematics and Evolution 98(2): 435-453. https://doi.org/10.3897/zse.98.86299

Lobón-Rovira J, Conradie W, Baptista NL, Vaz Pinto P. 2022. A new species of feather-tailed leaf-toed gecko, Kolekanos Heinicke, Daza, Greenbaum, Jackman, Bauer, 2014 (Squamata, Gekkonidae) from the poorly explored savannah of western Angola. ZooKeys 1127: 91-116.https://doi.org/10.3897/zookeys.1127.84942

Conradie W, Keates C, Baptista NL, Lobón-Rovira J. 2022. Taxonomical review of Prosymna angolensis Boulenger, 1915 (Elapoidea, Prosymnidae) with the description of two new species. ZooKeys 1121: 97–143. https://doi.org/10.3897/zookeys.1121.85693  

Reeves B, Brooke CF, Venter JA, Conradie W. 2022. The Reptiles and Amphibians of the Mpofu-Fort Fordyce Nature Reserve Complex in the Winterberg Mountains, Eastern Cape Province, South Africa African Journal of Wildlife Research 52(1), (12 September 2022). https://doi.org/10.3957/056.052.0134

Marques MP, Ceríaco LMP, Heinicke MP, Chehouri RM, Conradie W, Tolley KA, Bauer AM. 2022. The Angolan bushveld lizards, genus Heliobolus Fitzinger, 1843 (Squamata: Lacertidae): Integrative taxonomy and the description of two new species. Vertebrate Zoology 72: 745–769. https://doi.org/10.3897/vz.72.e85269

Farooq H, Nanvonamuquitxo C, Nassongole B, Conradie W, Bills R, Soares A, Antonelli A. 2022. Shedding Light on a Biodiversity Dark Spot: Survey Of Amphibians and Reptiles of Pemba Region in Northern Mozambique. Herpetological Conservation and Biology 17(2):423–432. http://www.herpconbio.org/Volume_17/Issue_2/Farooq_etal_2022.pdf 

Keates C, Conradie W, Dalu T, Dondofema F, Riddell ES, Wasserman RJ. 2022. Phylogenetic placement of the enigmatic Floodplain water snake, Lycodonomorphus obscuriventris FitzSimons, 1964. Koedoe 64(1): a1698. https://doi.org/10.4102/koedoe.v64i1.1698  

Channing A, Schmitz A, Zancolli G, Conradie W, Rödel M.-O. 2022. Phylogeny and taxonomy of the African frog genus Strongylopus (Anura: Pyxicephalidae). Revue Suisse de Zoologie 129: 243–281. https://doi.org/10.35929/RSZ.0074   

Telford N, Alexander GJ, Becker FS, Conradie W, Jordaan A, Kemp L, Le Grange A, Rebelo AR, Strauss S, Taft JM, Weeber J, Tolley KA. 2022. Extensions to the known geographic distributions of reptiles in the Karoo, South Africa. Herpetological Conservation and Biology 17(1):145–154. http://www.herpconbio.org/Volume_17/Issue_1/Telford_etal_2022.pdf

Lobón-Rovira J, Conradie W, Vaz Pinto P, Keates, C, Edwards, S, Du Plessis, Branch WR. 2022. Systematic revision of Afrogecko ansorgii (Boulenger, 1907) (Sauria: Gekkonidae) from western Angola. Zootaxa 5124 (4): 401–430. https://doi.org/10.11646/zootaxa.5124.4.1 

Lobón-Rovira J, Vaz Pinto P, Becker FS, Tolley KA, Measey J, Bennet B, Boon B, Sá S, Conradie W. 2022. An updated herpetofaunal species inventory of Iona National Park in southwestern Angola. Check List 18 (2): 289–321. https://doi.org/10.15560/18.2.289

Busschau T, Jordaan A, Conradie W, Daniels SR. 2022. Pseudocongruent phylogeography reflects unique responses to environmental perturbations in a biodiversity hotspot. Journal of Biogeography, 49(3): 1– 15. https://doi.org/10.1111/jbi.14334

2021

Baptista NL, Vaz Pinto P, Keates C, Edwards S, Roedel M-O, Conradie W. 2021. A new species of red toad, Schismaderma Smith, 1849 (Anura: Bufonidae), from central Angola. Zootaxa. 5081: 301–332. https://doi.org/10.11646/zootaxa.5081.3.1

Tolley KA, Nelford NS, Taft JM, Bates MF, Conradie W, Makhubo BG, Alexander GJ. 2021. Taxonomic inflation due to inadequate sampling: are girdled lizards (Cordylus minor species complex) from the Great Karoo one and the same? Biological Journal of the Linnean Society, blab119, https://doi.org/10.1093/biolinnean/blab119

Conradie W, Baptista NL, Verburgt L, Keates C, Harvey J, Júlio T, Neef G. 2021. Contributions to the herpetofauna of the Angolan Okavango-Cuando-Zambezi river drainages. Part 1: Serpentes (snakes). Amphibian & Reptile Conservation 15(2) [General Section]: 244–278 (e292). http://amphibian-reptile-conservation.org/pdfs/Volume/Vol_15_no_2/ARC_15_2_[General_Section]_244-278_e292.pdf

Alexander GJ, Tolley KA, Maritz B, McKechnie A, Manger P, Thomson RL,...Conradie W,.... et al. 2021. Excessive red tape is strangling biodiversity research in South Africa. S Afr J Sci. 2021;117(9/10), Art. #10787. https://doi.org/10.17159/sajs.2021/10787

Vaz Pinto P, Conradie W, Becker FS, Lobón-Rovira J. 2021. Updated distribution of Kolekanos plumicaudus (Sauria: Gekkonidae), with some comments on its natural history. Hepetoology Notes volume 14: 1207-1212. https://www.biotaxa.org/hn/article/view/69898/68437 

Parrinha D, Marques MP, Heinicke MP, Khalid F, Parker KL, Tolley KA, Childers JL, Conradie W, Bauer AM, Ceríaco LM P. 2021. A revision of Angolan species in the genus Pedioplanis Fitzinger (Squamata: Lacertidae), with the description of a new species. Zootaxa 5032 (1): 001–046 https://doi.org/10.11646/zootaxa.5032.1.1 

Conradie W, Ferreira R, Vaz Pinto P, Lobón-Rovira, J. 2021. Updated geographic distribution of Rhamnophis aethiopissa (Squamata: Colubridae) for Angola, with comments on diet. Herpetology Notes, volume 14: 1105-1109. https://www.biotaxa.org/hn/article/view/68514/68178 

Reeves B, Mdoko S, Rebelo A.D, Conradie W. 2021. Herpetological survey of The Island Nature Reserve in Nelson Mandela Bay Municipality, Eastern Cape Province, South Africa. Herpetology Notes, volume 14: 1087-1099. https://www.biotaxa.org/hn/article/view/67441/68070

Lobón-Rovira J, Conradie W, Buckley Iglesias D, Ernst R, Verisimmo L, Baptista N, Vaz Pinto P 2021 Between sand, rocks and branches: an integrative taxonomic revision of Angolan Hemidactylus Goldfuss, 1820, with description of four new species. Vertebrate Zoology 71: 465-501. https://doi.org/10.3897/vz.71.e64781

David G. Chapple, Uri Roll, Monika Böhm, Rocío Aguilar, Andrew P. Amey, Chris C. Austin, Marleen Baling, Anthony J. Barley, Michael F. Bates, Aaron M. Bauer, Daniel G. Blackburn, Phil Bowles, Rafe M. Brown, S.R. Chandramouli, Laurent Chirio, Hal Cogger, Guarino R. Colli, Werner Conradie,....Shai Meiri. 2021. Conservation status of the world's skinks (Scincidae): Taxonomic and geographic patterns in extinction risk, Biological Conservation, 257 (2021) 109101. https://doi.org/10.1016/j.biocon.2021.109101. 

Deepak, V, Maddock, ST,  Williams, R, Nagy, ZT, Conradie, W,  Rocha, S, Harris, DJ,  Perera, A, Gvoždík, V, Doherty-Bone, TM, Kamei, Rachunliu G, Menegon, M, Labisko, J,  Morel, C, Cooper, N, Day, JJ,  Gower. DJ. 2021 Molecular phylogenetics of sub-Saharan African natricine snakes, and the biogeographic origins of the Seychelles endemic Lycognathophis seychellensis, Molecular Phylogenetics and Evolution, 2021,107152, ISSN 1055-7903, https://doi.org/10.1016/j.ympev.2021.107152.

Nicolau, GK, Petford, M, Edwards, S, Busschau, T,Lynch, K, Kemp, L, Balmer, J, Keates, C, Hundermark, CR, Weeber, J, Conradie, W. 2021. New insights into the geographical distribution, ecology and conservation status of South Africa’s endemic Coastal Leaf-toed Gecko, Cryptactites peringueyi (Boulenger, 1910). Herpetology Notes, volume 14: 439-450 https://www.biotaxa.org/hn/article/view/63908/65534

Conradie, W & Vaz Pi tnto, P. 2021. A snake with an appetite for the rare: Amblyodipsas polylepis (Bocage, 1873) feeding on the amphisbaenid Monopeltis luandae Gans, 1976.  Herpetology Notes 14: 205-207 https://www.biotaxa.org/hn/article/view/64415 

Branch WR, Schmitz A, Lobón-Rovira J, Baptista NL, António T, Conradie W 2021. Rock island melody: A revision of the Afroedura bogerti Loveridge, 1944 group, with descriptions of four new endemic species from Angola. Zoosystematics and Evolution 97(1): 55-82. https://doi.org/10.3897/zse.97.57202

2020

Conradie W, Keates C, Lobón-Rovira J, Vaz Pinto P, Verburgt L, Baptista NL, Harvey J, Júlio T. 2020. New insights into the taxonomic status, distribution and natural history of De Witte’s Clicking Frog (Kassinula wittei Laurent, 1940), African Zoology, DOI: 10.1080/15627020.2020.1821771

Nielsen SV, Conradie W, Ceríaco LMP, Bauer AM, Heinicke MP, Stanley EL, Blackburn DC 2020. A new species of Rain Frog (Brevicipitidae, Breviceps) endemic to Angola. ZooKeys 979: 133–160. https://doi.org/10.3897/zookeys.979.56863

Conradie W, Reeves B, Mdoko S, Pamla L, Gxabhu, O. 2020. Herpetofaunal survey of the Ongeluksnek (Malekgalonyane) Nature Reserve in the foothills of the Drakensberg, Eastern Cape Province, South Africa. Herpetology Notes, volume 13: 717-730 https://www.biotaxa.org/hn/article/view/61297/62930 

Bittencourt-Silva GB, Bayliss J, Conradie W. 2020. First herpetological surveys of Mount Lico and Mount Socone, Mozambique. Amphibian & Reptile Conservation 14(2) [General Section]: 198–217 (e247). http://amphibian-reptile-conservation.org/pdfs/Volume/Vol_14_no_2/ARC_14_2_[General_Section]_198-217_e247.pdf 

Busschau T, Conradie W, Daniels SR. 2020. One species hides many: Molecular and morphological evidence for cryptic speciation in a thread snake (Leptotyphlopidae: Leptotyphlops sylvicolus Broadley & Wallach, 1997). J Zool Syst Evol Res. 2020;00:1–27. https://doi.org/10.1111/jzs.12401  

Hallermann J, Ceríaco LMP, Schmitz A, Ernst R, Conradie W, Verburgt L, Marques MP,  Bauer AM. 2020. A review of the Angolan House snakes, genus Boaedon Duméril, Bibron and Duméril (1854) (Serpentes: Lamprophiidae), with description of three new species in the Boaedon fuliginosus (Boie, 1827) species complex, African Journal of Herpetology, DOI: 10.1080/21564574.2020.1777470

Conradie W, V Deepak, Keates C,  Gower DJ. 2020. Kissing cousins: a review of the African genus Limnophis Günther, 1865 (Colubridae: Natricinae), with the description of a new species from north-eastern Angola, African Journal of Herpetology, DOI: 10.1080/21564574.2020.1782483

Kushata JNT, Conradie W, Cherry MI, Daniels SR. 2020. Comparison of the mitochondrial phylogeographical structure of a generalist and two specialist frog species reveals contrasting patterns in the Eastern and Western Cape provinces of South Africa, Biological Journal of the Linnean Society, blaa049, https://doi.org/10.1093/biolinnean/blaa049

Conradie W, Busschau T, Jordaan A. 2020. Do Nucras livida occur along the West Coast of South Africa? A review of recently collected and historical material. Herpetology Notes 13:19-24. https://www.biotaxa.org/hn/article/view/57362/59391 

Engelbrecht HM, Branch WR, Greenbaum E, Burger M, Conradie W, Tolley KA. 2020. African Herald snakes, Crotaphopeltis, show population structure for a widespread generalist but deep genetic divergence for forest specialists. J Zool Syst Evol Res. 2020;00:1–14. https://doi.org/10.1111/jzs.12361  

2019

Kilunda FK, Conradie W, Wasonga DV, Jin J-Q, Peng M-S, Murphy RW, Malonza PK, Che J. 2019. Revalidation and resurrection of Panaspis massaiensis (Angel, 1924) and the description of a new species of Panaspis Cope (Squamata: Scincidae) from south-eastern Kenya. Zootaxa 4706 (2): 255–274. https://doi.org/10.11646/zootaxa.4706.2.3 

Keates C, Conradie W, Greenbaum E, Edwards S. A snake in the grass: Genetic structuring of the widespread African grass snake (Psammophylax Fitzinger 1843), with the description of a new genus and a new species. J Zool Syst Evol Res. 2019;57:1039–1066. https://doi.org/10.1111/jzs.12337      

Maritz R, Conradie W, Sardinha CI, Peto A, Chechene AHD, Maritz B. 2019. Ophiophagy and cannibalism in African vine snakes (Colubridae: Thelotornis). Afr J Ecol. 2019;00:1–5. https://doi.org/10.1111/aje.12702  

Ihlow F, Farooq HM, Gvoždík V, Hofmeyr MD, Conradie W, Campbell PD, Harvey J, Verburgt L, Fritz U. 2019. Geographic range extension of Speke’s Hinge-back Tortoise Kinixys spekii Gray, 1863. Amphibian & Reptile Conservation 13(2) [Special Section]: 61–67 (e195).

Branch WR, Conradie W, Vaz Pinto P, Tolley KA. 2019. Another Angolan Namib endemic species: a new Nucras Gray, 1838 (Squamata: Lacertidae) from south-western Angola. Amphibian & Reptile Conservation 13(2) [Special Section]: 82–95 (e199)

Measey J, Tarrant J, Rebelo A, Turner A, Du Preez L, Mokhatla M, Conradie W. 2019. Has strategic planning made a difference to amphibian conservation research in South Africa? Bothalia 49(1), a2428. https://doi.org/10.4102/abc.v49i1.2428 

Conradie W, Grieneisen ML, Hassapakis CL (Editors). 2019. Compilation of personal tributes to William Roy Branch (1946–2018): a loving husband and father, a good friend, and a mentor. Amphibian & Reptile Conservation 13(2) [Special Section]: i–xxix (e186). http://amphibian-reptile-conservation.org/pdfs/Volume/Vol_13_no_2/ARC_13_2_[Special_Section]_i-xxix_e186_low_res.pdf

Branch WR, Bittencourt-Silva G, Conradie W, Engelbrecht E, Loader S, Menegon M, Nanvonamuquitxo C, Tolley T. 2019. A new species of tree snake (Dipsadoboa, Serpentes: Colubridae) from ‘sky island’ forests in northern Mozambique, with notes on other members of the Dipsadoboa werneri group. Zootaxa 4646 (3): 541–563. https://doi.org/10.11646/zootaxa.4646.3.6

Busschau T, Conradie W, Daniels S. 2019. Evidence for cryptic diversification in a rupicolous forest-dwelling gecko (Gekkonidae: Afroedura pondolia) from a biodiversity hotspot. Molecular Phylogenetics and Evolution 139, 106549 https://doi.org/10.1016/j.ympev.2019.106549 

Rebelo AD, Bates MF, Burger M, Branch WR, Conradie W. 2019. Range expansion of the Common Dwarf Gecko, Lygodactylus capensis: South Africa’s most successful reptile invader. Herpetology Notes, volume 12: 643-650. https://biotaxa.org/hn/article/view/46230/47319  

Portik DM, Bell RC, Blackburn DC, Bauer AM, Barratt CD, Branch WR, Burger M, Channing A, Colston TJ, Conradie W, et al. 2019. Sexual Dichromatism Drives Diversification within a Major Radiation of African Amphibians. Systematic Biology, syz023, https://doi.org/10.1093/sysbio/syz023

Portillo F, Stanley EL, Branch WR, Conradie W, Rödel M-O, Penner J, et al. 2019. Evolutionary history of burrowing asps (Lamprophiidae: Atractaspidinae) with emphasis on fang evolution and prey selection. PLoS ONE 14(4): e0214889. https://doi.org/10.1371/journal.pone.0214889  

Weinell JL, Branch WR, Colston TJ, Jackman TR, Kuhn A, Conradie W, Bauer AM. 2019. A species-level phylogeny of Trachylepis (Scincidae: Mabuyinae) provides insight into their reproductive mode evolution, Molecular Phylogenetics and Evolution (2019), doi: https://doi.org/10.1016/j.ympev.2019.04.002

Conradie W, Branch WR, Watson G. 2019. Type specimens in the Port Elizabeth Museum, South Africa, including the historically important Albany Museum collection. Part 2: Reptiles (Squamata). Zootaxa 4576 (1): 001 045. https://doi.org/10.11646/zootaxa.4576.1.1

Tolley KA, Weeber W, Maritz B, Verburgt L, Bates MF, Conradie W, Hofmeyr M, Turner AT, da Silva J, Alexander G. 2019. No safe haven: Protection levels show imperilled South African reptiles not sufficiently safe-guarded despite low average extinction risk. Biological Conservation 233 (2019) 61–72. https://doi.org/10.1016/j.biocon.2019.02.006 

Baptista N, Conradie W. Vaz Pinto P, Branch WR 2019. The Amphibians of Angola: Early Studies and the Current State of Knowledge. Pp 243–281. In: Biodiversity of Angola. Science & Conservation: A Modern Synthe­sis. Huntley BJ, Ferrand N, Russo V, Lages F. Springer, Switzerland. 552 p. https://rd.springer.com/chapter/10.1007/978-3-030-03083-4_12 

Branch WR, Baptista N, Vaz Pinto P, Conradie W. 2019. The Reptiles of Angola – History, updated checklists, endemism, hot spots, and future directions for re­search. Pp 283–326. In: Biodiversity of Angola. Science & Conservation: A Modern Synthe­sis. Huntley BJ, Ferrand N, Russo V, Lages F. Springer, Switzerland. 552 p. https://rd.springer.com/chapter/10.1007/978-3-030-03083-4_13 

Zhao Z, Verdú-Ricoy J, Mohlakoana S, Jordaan A, Conradie W, Heideman N. 2019. Unexpected phylogenetic relationships within the world's largest limbless skink species (Acontias plumbeus) highlight the need for a review of the taxonomic status of Acontias poecilus. Journal of Zoological Systematics and Evolutionary Research 2019;00:1–16. https://doi.org/10.1111/jzs.12263

Branch WR, Verburgt L, Bayliss J, Kucharzewski C, Rodel M-O, Conradie W. 2019. New records of the Large-eyed Green Snake, Philothamnus macrops (Boulenger 1895), from Mozambique. Herpetology Notes, 12: 19-29. [PDF] 

Engelbrecht HM, Branch WR, Greenbaum E, Alexander GJ, Jackson K, Burger M, Conradie W, Kusamba C, Zassi-Boulou, A-G, Tolley KA. 2019. Diversifying into the branches: species boundaries in African green and bush snakes, Philothamnus (Serpentes: Colubridae). Molecular phylogenetics and evolution 130: 357-365 https://doi.org/10.1016/j.ympev.2018.10.023

2018

Tolley KA, Conradie W, Harvey J, Measey J, Blackburn DG. 2018 Molecular phylogenetics reveals a complex history underlying cryptic diversity in the Bush Squeaker Frog (Arthroleptis wahlbergii) in southern Africa. African Zoology, 1-16 DOI: 10.1080/15627020.2018.1517608

Nielsen SV, Daniels SR, Conradie W, Heinicke MP, Noonan BP. 2018. Multilocus phylogenetics in a widespread African anuran lineage (Brevicipitidae: Breviceps) reveals patterns of diversity reflecting geoclimatic change. J Biogeogr. 2018;00:1–13. https://doi.org/10.1111/jbi.13394

Conradie W, Busschau T,  Edwards S. 2018. Two new species of Acontias form the Mpumalanga Highveld escarpment of South Africa. Zootaxa 4429(1):089-106. https://doi.org/10.11646/zootaxa.4429.1.3

Portillo F, Branch WR, Conradie W, Rödel M-O, Penner J, Bare, MF, Kusamba C, Muninga WM, Aristote MM, Bauer AM, Trape J-F, Nagy ZT, Carlino P, Pauwels OSG, Menegon M, Burger M, Mazuch T, Jackson K, Hughes DF, Behangana M, Greenbaum E. 2018. Phylogeny and biogeography of the African burrowing snake subfamily Aparallactinae (Squamata: Lamprophiidae), Molecular Phylogenetics and Evolution 127 (2018):288-303. doi: https://doi.org/10.1016/j.ympev.2018.03.019

Broadley, D.G., Tolley, A., Conradie, W., Wishart, S., Trape, J-F., Burger, M., Kusamba, C.,  Zassi-Boulou, A-G. & Greenbaum, E. 2018.  A phylogeny and genus-level revision of the African file snakes Gonionotophis Boulenger (Squamata: Lamprophiidae), African Journal of Herpetology, 67:1, 43-60, DOI: 10.1080/21564574.2018.1423578 

Conradie, W., Bittencourt-Silva, G.B., Farooq, H.M., Loader, S.P., Menegon, M. & Tolley K.A. 2018. New species of Mongrel Frogs (Pyxicephalidae: Nothophryne) for northern Mozambique inselbergs, African Journal of Herpetology, 67:1, 61-85, DOI: 10.1080/21564574.2017.1376714

Conradie, W., Verburgt, L., Portik, D.M., Ohler, A., Bwong, B.A. & Lawson, L.P. 2018. A new Reed Frog (Hyperoliidae: Hyperolius) from coastal northeastern Mozambique. Zootaxa, 4379: 177-198, http://dx.doi.org/10.11646/zootaxa.4379.2.2

Jongsma, G.F.M. ,  Barej, M.F. Barratt, C.D., Burger, M., Conradie, W., Ernst, R., Greenbaum, E., Hirschfeld, M.,  Leaché, A.D., Penner, J.,  Portik, D.M., Zassi-Boulou, A., Rödel, MO., &  Blackburn, D.C. 2018. Diversity and biogeography of frogs in the genus Amnirana (Anura: Ranidae) across sub-Saharan Africa, Molecular Phylogenetics and Evolution, Volume 120, March 2018, Pages 274-285, ISSN 1055-7903, https://doi.org/10.1016/j.ympev.2017.12.006

2017

Pietersen, D.W., Pietersen, E.W., & Conradie, W.  2017. Preliminary herpetological survey of the Ngonye Falls and surrounding regions in south-western Zambia. Amphibian & Reptile Conservation 11(1) [Special Section]: 24–43 (e148).[PDF]

Conradie, W & Bills, R. 2017. Wannabe Ranid: Notes on the morphology and natural history of the Lemaire’s Toad (Bufonidae: Sclerophrys lemairii). Salamandra 53(3):439-444. [PDF]

Branch, W.R., Haacke, H., Vaz Pinto, P., Conradie, W., Baptista, N., Verburgt, V., & Verisimmo, L. 2017. Loveridge’s Angolan geckos, Afroedura karroica bogerti and Pachydactylus scutatus angolensis (Sauria, Gekkonidae): new distribution records, comments on type localities and taxonomic status. Zoosystematics and Evolution, 93: 157-166. [PDF]

Busschau T, Conradie W, Jordaan A, Daniels SR. 2017 Unmasking evolutionary diversity among two closely related South African legless skink species (Acontinae: Acontias) using molecular data. Zoology 121:72-82. dx.doi.org/10.1016/j.zool.2016.11.005

2016

Conradie, W., Reeves, B., Brown, N. & Venter, J.A. 2016. Herpetofauna of the Oviston, Commando Drift and Tsolwana nature reserves in the arid interior of the Eastern Cape Province, South Africa. Indago 32:81-98 [PDF]

Ceríaco et al. 2016. Photography-based taxonomy is inadequate, unnecessary, and potentially harmful for biological sciences. Zootaxa 4196 (3): 435–445. [PDF]

Conradie, W., Bill, R., & Branch, W.R. 2016. The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola.  Amphibian & Reptile Conservation 10(2) [Special Section]: 6–36 (e126). [PDF]

Conradie, W., Bittencourt-Silva, Engelbrecht, H.M., Loader, S. P., Menegon, M., Nanvonamuquitxo, C., Scott, M. & Tolley, K.A. 2016. Exploration into the hidden world of Mozambique’s sky island forests: new discoveries of reptiles and amphibians. Zoosystematics and Evolution, 92 (2): 163-180. [PDF]

Metallinou, M., Weinell, J.L., Karin, B. R., Conradie, W., Wagner, P., Schmitz, A., Jackman, T. R. & Bauer, A. 2016. A single origin of extreme matrotrophy in African mabyine skinks. Biology Letters, 12: 20160430. [PDF]

Medina, M.F, Bauer, A.M., Branch, W.R., Schmitz, A., Conradie, W., Nagy, Z.T., Hibbitts, T.J., Ernst, R., Portik, D.M., Nielsen, S.V., Colston, T.J., Kusamba, C., Behangana, M., Rödel, M-O. & Greenbaum, E. 2016. Molecular phylogeny of Panaspis and Afroablepharus skinks (Squamata: Scincidae) in the savannas of sub-Saharan Africa, Molecular Phylogenetics and Evolution, 100:409-423. [PDF]

Poynton, J.C., Loader, S.P., Conradie, W., Rodel, M-O &Liedtke, C. 2016. Designation and description of a neotype of Sclerophrys maculata (Hallowell, 1854), and reinstatement of S. pusilla (Mertens, 1937) (Amphibia: Anura: Bufonidae). Zootaxa 4089: 73-94. [PDF]

Conradie, W; Bittencourt-Silva, G., Loader, S.P., Menegon, M., Nanvonamuquitix, X., Kotze, A., Dalton, D.L., Engelbrecht, H.M. & Tolley, K.A. 2016. Batrachochytrium dendrobatidis survey of amphibians in the northern Mozambique ‘Sky Islands’ and low lying areas. Herpetological Review 47: 42-46 [PDF]

Bittencourt-Silva, G., Conradie, W., Siu-Ting, K., Tolley, K.A., Channing, A., Cunningham, M., Farooq, H.M., Menegon, M. & Loader, S.P. 2016. The phylogenetic position and diversity of the enigmatic mongrel frog Nothophryne Poynton, 1963 (Amphibia, Anura). Molecular Phylogenies and Evolution 99: 89-102.[PDF]

2015

Farooq, H., Liedtke, H. C., Bittencourt-Silva, G., Conradie, W., & Loader, S. 2015. The distribution of Mertensophryne anotis with a new record in Northern Mozambique. Herpetology Notes 8: 305-307. [PDF]

Farooq, H.O.M. & Conradie, W. 2015. A second record of Scolecomorphus kirkii Boulenger, 1883 (Gymnophiona: Scolecomorphidae) for Mozambique. Herpetology Notes, 8: 59-62. [PDF]

Venter, A.J. and Conradie, W. 2015. A checklist of the reptiles and amphibians found in the protected areas along the South African Wild Coast, with notes on conservation implications. Koedoe 57 (1): 1-25. http://dx.doi.org/10.4102/koedoe.v57i1.1247 

Conradie, W. and Conradie, C. 2015. Correlation between development and increase of number of labial tooth rows in Ghost Frog tadpoles (Anura: Heleophrynidae). Acta Herpetologica 10(2): 143-148. [PDF]

Conradie, W, Branch, W. R. & Watson, G. 2015. Type specimens in the Port Elizabeth Museum, South Africa, including the historical important AlbanyMuseum collection. Part 1: Amphibians. Zootaxa 3936 (1):042-070 [PDF]

2014

Conradie, W. 2014. The King of the Dwarves: a new cryptic species of Dainty Frog (Anura: Pyxicephalidae: Cacosternum) from the eastern Great Escarpment of South Africa. Zootaxa 3785 (3): 438-452.  http://dx.doi.org/10.11646/zootaxa.3785.3.6 

2013

Conradie, W., Branch, W.R., and Tolley, K.A. 2013. Fifty Shades of Grey: giving colour to the poorly known Angolan Ashy reed frog (Hyperoliidae: Hyperolius cinereus), with the description of a new species. Zootaxa 3635 (3): 201-223.

Channing, A., Hillers, A., Lötters, S., Rödel, M.O., Schick, S., Conradie, W, Rödder, R., Mercurio, V., Wagner, P., Dehling, M.J., Du Preez, L.H., Kielgast, J. & Burger, M. 2013. Taxonomy of the super-cryptic Hyperolius nasutus group of Long reed frogs of Africa (Anura: Hyperoliidae), with descriptions of six new species. Zootaxa 3620 (3): 301-350.

2012

Conradie, W., Branch, W.R., Measey, J.G. and Tolley, K.A. 2012. A new species of Hyperolius Rapp, 1842 (Anura: Hyperoliidae) from the Serra da Chela mountains, south-western Angola. Zootaxa3269: 1–17.

Conradie, W., Measey, J.G., Branch, W.R. and Tolley, K.A. 2012. Revised phylogeny of African sand lizards (Pedioplanis), with the description of two new species. African Journal of Herpetology 61: 91-112.

Conradie, W., Tweddle, D., Makinen, T. & Bills, R. 2012 Description of the Hylarana darlingi (Anura: Ranidae) from the Mulanje Area, Malawi. Zootaxa3441:56-58.

2011

Conradie, W., Weldon, C., Smith, K.G. & Du Preez, L.H. 2011. Seasonal pattern of chytridiomycosis in common river frog (Amietia angolensis) tadpoles in the South African Grassland Biome. African Zoology 46(1): 95–102.

Conradie, W., Harvey, J., Kotze, A., Dalton, D.L. and Cunningham, M.J. 2011. Confirmed amphibian chytrid in Mount Mulanje Area, Malawi. Herpetological Review 43(3): 369-371.

2010

Conradie, W., Branch, W.R., Braack, H. and Mason, M. 2010. Notes on the diet of recently metamorphosed Giant African Bullfrogs (Anura: Pyxicephalidae: Pyxicephalus adspersus) and growth increase during the first nine months in a semi-natural habitat. Herpetology Notes, volume 3: 215-219.

2007

Smith, K.G, Weldon, C., Conradie, W & Du Preez, L.H. 2007. Relationships among size, development, and Batrachochytrium dendrobatidis infection in African tadpoles. Diseases of Aquatic Organisms 74: 159-164.

 

Books/Book Chapters

Tolley KA, Conradie W, Pietersen DW, Weeber J, Burger M, Alexander, G.J. (eds) 2023. Conservation status of the reptiles of South Africa, Eswatini and Lesotho. Suricata 10. South African National Biodiversity Institute, Pretoria

Baptista N, Conradie W. Vaz Pinto P, Branch WR 2019. The Amphibians of Angola: Early Studies and the Current State of Knowledge. Pp 243–281. In: Biodiversity of Angola. Science & Conservation: A Modern Synthe­sis. Huntley BJ, Ferrand N, Russo V, Lages F. Springer, Switzerland. 552 p. 

Branch WR, Baptista N, Vaz Pinto P, Conradie W. 2019. The Reptiles of Angola – History, updated checklists, endemism, hot spots, and future directions for re­search. Pp 283–326. In: Biodiversity of Angola. Science & Conservation: A Modern Synthe­sis. Huntley BJ, Ferrand N, Russo V, Lages F. Springer, Switzerland. 552 p. 

Branch, W.R. & Conradie, W. 2011.  Herpetofauna of the Carumba Lagoon Area, pp195-209. In: Huntley, B. (ed.), Expedition Report. Rapid Biodiversity assessment of the Carumba Lagoon area, Lunda Norte, Angola. Ministerio do Ambiente de Angola, 219p

Conradie, W. & Morrison, C. 2011. Amphibians In. Our Coastal Treasure: The Greater St Francis Area.

Conradie, W., Smith, K.G., Weldon, C., & Du Preez, L.H. 2006. Field Guide to the Frogs and Toads of the Vredefort Dome World Heritage Site. Platinum Press. Potchefstroom.

Shorter Communications

Conradie W, Booysen M. 2025. Geographical Distribution: Hemirhagerrhis notaenia. African Herp News 88: 23–35

Conradie E, Botma D, Nanvonamuquito C. 2025. Geographical Distribution: Rieppeleon brachyrus. African Herp News 88: 29–34

Conradie, W., Nicolau, G.K., Kemp, L., Edwards, S., Klein Snakenborg, S., Reeves, B. 2020. Geogrpahical Distributions: Lygodactylus capensis. African Herp News 74: 78–82

Miguel, A.R. & Conradie, W. 2019. Geographical Distribution: Hyperolius stictus Conradie, Verburgt, Ohler, Bwong, Lawson, 2018. African Herp News 71:44–46.

Nicolau, G.K., Kemp, L. & Conradie, W. 2018. Geographical Distribution: Panaspis wahlbergii Smith, 1849. African Herp News 69:27–30.

Rebelo, A.R, Zhao, Z, Jordaan, A., Jordaan. P.R. & Conradie, W. 2018. Geographical Distribution Scelotes caffer Peters, 1861. African Herp News 68:53–56.

Rebelo, A.R, Jordaan. P.R. & Conradie, W. 2018. Geographical Distribution Ptenopus garralus maculatus Gray, 1866. African Herp News 68:50–52.

Conradie, W & Herbert, D.G. 2018. The story of the snail and the gecko egg. Biodiversity Observations 9.3:1-2

Conradie, W. & Busschau, T. Geographical Distribution: Macrelaps microlepidotus. African Herp News 67:41-43.

Kemp, L. & Conradie, W. Geographical Distribution: Aspidelaps sulcatus. African Herp News, 65:42-45.

Branch, W.R  & W. Conradie. 2017 Phrynomantis affinis Maximum Size. African Herp New, 65: 12-14.

Conradie, W., Venter, J & Du Preez, B. 2017. Geographical Distribution Note: Tropidosaura montana natalensis. African Herp News, 65:40-42

Conradie, W., Branch W.R. & Herbert, H.. 2017 Tetradactylus seps Diet. African Herp News, 65:14-16

Conradie W., King, F.V., Pressley M., & Leander, A. 2017. Natural History: Gerrhosaurus typicus. African Herp News 64:28-30

Conradie, W; Bittencourt-Silva, G., Loader, S.P., Menegon, M., Nanvonamuquitix, X., Kotze, A., Dalton, D.L., Engelbrecght, H.M. & Tolley, K.A. 2017. Batrachochytrium dendrobatidis survey of amphibians in the northern Mozambique ‘Sky Islands’ and low lying areas. FrogLog 118:95-96

Branch, W.R & Conradie, W. 2013. Naja (Boulengerina) annulata annulata - Geographical Distribution Diet. African Herp News 59:50-53

Conradie, W & Bourquin, S. 2013. Geographical Distributions: Acontias kgalagadi kgalagadi (Lamb, Biswas & Bauer, 2010): African Herp News: 60:29-30.

Conradie, W. 2012. Pedioplanis husabensis: Maximum Size. Herpetological Bulletin 120: 42

Conradie, W. 2012. Pedioplanis lineoocellata lineoocellata: Maximum Size. Herpetological Bulletin 120: 42-43

Conradie, W. 2012. Varanus albigularis - Diet. African Herp News 58:15-16

Conradie, W., Venter, J. & Nicolau, J.R. 2012. Geographical Distribution – Macrelaps microlepidotus. African Herp News 58:19-21

Conradie, W. 2012. Geographical Distribution – Nucras taeniolata. African Herp News 58: 21-22

Conradie, W. & Venter Jan. 2012. Hluleka Nature Reserve: unexpected amphibian diversity. FrogLog 103:22-23.

Conradie, W.  2012. First new frog discovery for Angola in 40 years. FrogLog 103:25.

Nielsen, S.V. and Conradie, W. 2011. Psammophis: Mortality. African Herp News 55:17-18

Conradie, W. and Tarrant, J. 2011. Amatola Toad AWOL: Thirteen years of futile searches. Froglog 97: 24

Conradie, W. and Weldon, C. 2011. Monitoring of the endangered Hewitt’s ghost frog. Froglog 97: 31.

Conradie, W., Bills, R and Bok, A. 2011. Leptopelis bocagii – Maximum Size. African Herp News 53: 42-43.

Conradie, W., Du Preez, L.H., Smith, K.G., & Weldon, C. 2008. Herpetological survey: Frogs of the Vredefort Dome Conservation Area. African Herp News 44:23-25